Cozy and her littermates are driven from their suburban den because something is killing the foxes in their area, and so they travel to a strange new world, the City, to confront a new set of dangers, where they meet 0-370, an injured fox who has escaped from a fur farm and who gives them a new mission--return to the farm, defeat the humans, and free the caged foxes.

Once best friends, Arthur and Wally find themselves on opposites sides of an epic war. Arthur, now officially a Novitiate of the Wardens of Weirdwood, is tasked with defending the border between the Real and Imaginary realms. But Wally and his brother Graham, who can see into the future, seek to topple it and create a new world. Meanwhile, the shadowy Order of Eldar, led by the fearsome Eraser, are also prepared to sow chaos and let wondrous and dangerous...

Phoebe, fifteen, the daughter of a famous mother and unearthly father, suddenly begins experiencing radical changes as she enters one scene after another from 1950s and 60s science fiction movies.